Warren D. Hauk, PhD

• Phylogeny of Ophioglossaceae based on morphological and molecular data
• Evolutionary relationships in Botrychium s.s., the moonworts
• Ancestry of allopolyploids in Botrychium s.s. using chloroplast and nuclear markers
• ISSR variation among populations of Sceptridium dissectum
• Genetic variation within and among North American species of Sceptridium, the grapeferns, using ISSR markers and DNA sequences
• Evolution of vascular trace structure in Ophioglossum s.s., the adder's tongue ferns

As a plant systematist my main research focus is the fern family Ophioglossaceae, commonly called the adder's tongue ferns. I use both molecular and morphological evidence to infer evolutionary patterns and processes within the family and to establish classifications that reflect evolutionary history. Currently my research centers on two main species groups, Botrychium s.s. (the moonworts) and Sceptridium (the evergreen grapeferns). In both species groups I use DNA sequencing and/or ISSR markers to address evolutionary relationships among species, the distribution of genetic variation within species, and the interface between species and populations.

• Associate Professor of Biology, Denison University:, 2004-present
• Assistant Professor of Biology, Denison University:, 1998-2004
• Postdoctoral Researcher, Missouri Botanical Garden, St. Louis, MO: 1996-1998
• Ph.D.University of North Carolina, Chapel Hill, NC: (biology)1996
• M.A.University of Kansas, Lawrence, KS; (botany) 1990
• B.A.Baylor University, Waco, TX; (biology, anthropology) 1985

• Biol 150 Introduction to the Science of Biology
• Biol 202 Ecology & Evolution
• Biol 320 Plant Systematics
• Biol 380 Advanced Evolutionary Biology

• Small, R.B., E.B. Lickey, J. Shaw, and W.D. Hauk. 2005. Amplification of non-coding chloroplast DNA for phylogenetic studies in Lycophytes and Pteridophytes with a comparative example of relative phylogenetic utility from Ophioglossaceae. Molecular Phylogenetics and Evolution 36: 509-522.
• Hauk, W. D., C. R. Parks, and M.W. Chase. 2003. Phylogenetic studies of Ophioglossaceae: Evidence from rbcL and trnL-F plastid DNA sequences and morphology. Molecular Phylogenetics and Evolution 28(1): 131-151.
• Hauk, W. D. and M. S. Barker. 2003. Botrychium lanceolatum subsp. angustisegmentum in Ohio. American Fern Journal 93(2): 93-94.
• Barker, M. S. and W. D. Hauk. 2003. An evaluation of Sceptridium dissectum (Ophioglossaceae) with ISSR markers: implications for Sceptridium systematics. American Fern Journal 93(1): 1-19.
• Hauk, W.D. In press. A review of the genus Memora (Bignoniaceae). Annals of the Missouri Botanical Garden.
• Hauk, W.D. 2002. Revisions of the genera Potamoganos and Roentgenia (Bignoniaceae). Annals of the Missouri Botanical Garden 89: 77-87.
• Hauk, W.D. and M.S. Barker. 2000. Larch Mountain, 2000 AFS Foray. Fiddlehead Forum: Bulletin of the American Fern Society 27(5): 26-27.
• Hauk, W.D. and C.H. Haufler. 1999. Isozyme variation and species relationships within the moonworts (Botrychium subgenus Botrychium). American Journal of Botany 86(5): 614-633.
• Hauk, W.D. 1999. Four new species of Memora (Bignoniaceae) from South America. Novon 9(1): 48-54.
• Hauk, W.D. 1998. A review of the genus Paragonia (Bignoniaceae). Annals of the Missouri Botanical Garden 85: 460-474.
• Hauk, W.D. 1998. A review of the genus Cydista (Bignoniaceae). Annals of the Missouri Botanical Garden 84: 815-840.
• Hauk, W.D. 1995. A molecular assessment of species relationships among cryptic species of Botrychium subgenus Botrychium (Ophioglossaceae). American Fern Journal 85(4): 375-394.
• Hasebe, M., P.G. Wolf, K.M. Pryer, K. Ueda, M. Ito, R. Sano, G.J. Gastony, J. Yokoyama, J.R. Manhart, N. Murakami, E.R. Crane, C.H. Haufler, and W.D. Hauk. 1995. Fern phylogeny based on rbcL nucleotide sequences. American Fern Journal 85(4): 134-181.

  Manuscripts in preparation or submitted for publication

• Hauk, W.D, L.F. Kennedy, and H.M. Hawke. Phylogenetic relationships among species of Botrychium s.s. (Ophioglossaceae) based on three plastid DNA sequence data sets.

Published and Submitted Abstracts

• Hauk, W. D. 2005. Phylogenetic studies of Ophioglossaceae based on analyses of three plastid DNA data sets and morphology. Botanical Society of America, Botany 2005.
• Hauk, W. D., K.B. Gibson, and T.C. Masters. 2004. An evaluation of genetic variation in Sceptridium biternatum and S. lunarioides using ISSR markers. Botanical Society of America, Botany 2004.
• Hauk, W. D., L. F. Kennedy and H.M. Hawke. 2003. Phylogenetic relationships among species of Botrychiums.s. (Ophioglossaceae) based on three plastid DNA sequence data sets. Botanical Society of America, Botany 2003.
• Kennedy, L. F., H. M. Hawke, and W. D. Hauk. 2003. A Phylogenetic Analysis of Cryptic Moonwort Species (Botrychium s.s.: Ophioglossaceae) Using rbcL and trnL-F cpDNA Sequences. Midwest Ecology and Evolution Conference. University of Akron, Akron, OH.
• Hauk, W. D. and H. M. Hawke. 2002. Phylogenetic relationships in Botrychium s.s. (Ophioglossaceae) based on rbcL and trnL-F cpDNA sequences. Botanical Society of America, Botany 2002.
• Hawke, H.M. (W.D. Hauk) 2001. A phylogenetic analysis of cryptic moonwort species (Botrychium s.s.: Ophioglossacae) using rbcL and trnL-F DNA sequences. To Ohio Academy of Sciences, submitted Nov. 1, 2001.
• Barker, M.S. and W.D. Hauk. 2001. An evaluation of Sceptridium dissectum and S. oneidense (Ophioglossacae) using ISSR markers: implications for Sceptridium species circumscriptions. Botanical Society of America, Botany 2001.
• Barker, M.S., and W.D. Hauk. 2001. "Genetic variation of Sceptridium dissectum (Ophioglossaceae) assessed by ISSR (Inter-Simple Sequence Repeat) markers." The Ohio Journal of Science 101(1): 43.
• Hauk, W.D. 2000. Phylogeny of the Ophioglossaceae based on molecular and morphological characters. American Journal of Botany 87(6): 90.
• Farrar, D.R., C.L. Johnson-Groh, and W.D. Hauk. 2000. Biology and conservation of the Ophioglossaceae: A tribute to Warren "Herb" Wagner. American Journal of Botany 87(6): 89.
• Barker, M.S. (W.D. Hauk) 2000: Genetic Variation in Botrychium dissectum Sprengel (Ophioglossaceae) Assessed by ISSR (Inter-Simple Sequence Repeat) Markers. The Ohio Journal of Science 100(1): A8.
• Barker, M.S. (W.D. Hauk) 2000: A molecular assessment of infraspecific genetic variation in Draba verna L. using ISSR (Inter-Simple Sequence Repeat) Markers. The Ohio Journal of Science 100(1): A37.
• Robie, A.C. and W.D. Hauk. 1999. Evolutionary relationships among species of tropical lianas: assembling a molecular phylogeny of the tribe Bignonieae (Bignoniaceae). The Ohio Journal of Science 99(1): A8.
• Hauk, W.D., and S.S. Renner. 1999. Phylogenetic relationships among genera of the liana tribe Bignonieae (Bignoniaceae). Published by the XVI International Botanical Congress, Missouri Botanical Garden Press.
• Hauk, W.D., L.G. Lohmann, and S.S. Renner. 1997. Indels of the trnL-F intergenic spacer as generic markers in woody tropical lianas (Bignoniaceae tribe Bignonieae). American Journal of Botany 84(6): 201.
• Hauk, W.D., C.R. Parks, and M. W. Chase. 1996. A comparison between trnL-F intergenic spacer and rbcL DNA sequence data: an example from Ophioglossaceae. American Journal of Botany 83(6): 126.
• Hauk, W.D. 1994. A molecular assessment of cryptic speciation in Botrychium subgenus Botrychium. American Journal of Botany (supplement). Vol. 81(6): 121.
• Hauk, W.D. and M.W. Chase. 1993. Molecular systematics of the Ophioglossaceae. American Journal of Botany (supplement) Vol. 80(6): 109.
• Hauk, W.D. and M. W. Chase. 1991. Gene sequence systematics of the Ophioglossaceae. American Journal Botany (supplement) Vol. 78(6): 153.
• Hauk, W.D., C.H. Haufler, and P. G. Gegenheimer. 1989. Systematics of the genus Botrychium: the molecular story. American Journal of Botany (supplement) Vol. 76(6): 204.